Paul Downing has joined Blackburn Rovers for an undisclosed fee.
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Defender Ben Tilney has joined Brackley Town on a season-long loan deal.
Robbie Neilson has further strengthened his attacking options with the signing of promising striker Ryan Seager from Southampton.
MK Dons have swooped to secure the services of Wolverhampton Wanderers defender Ethan Ebanks-Landell on a season-long loan deal.
Ebanks-Landell began his career with Wolverhampton Wanderers, progressing through the Academy set-up at Molineux before turning professional in December 2010.
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